Why Down Wood Percent Cover?

For DecAID, we chose to describe amounts of down wood used by wildlife as percent cover because it is a good metric to describe the abundance of down wood as it relates to wildlife use (Carey 1995, Carey and Johnson 1995). For many wildlife species down wood provides cover. In our literature review, we found that percent cover was the most common measure used in studies showing wildlife selection of areas with higher amounts of down wood. Volume or biomass may be more relevant measures of ecosystem processes (e.g., nutrient cycling, site productivity, carbon sequestration, fuel loading), but we are opting to emphasize wildlife habitat.

Percent cover is one of the most precise and efficient means of recording down wood (Bate et al. 2002).By laying out a series of randomly placed line transects and measuring the length of the transect that intersects down wood, one can quickly and accurately determine the percent cover. The down wood falling along the transect can be categorized by diameter, species or decay class, further describing the characteristics of the down wood in the area of interest. In contrast, estimates of volume, biomass, pieces per acre, or total length require more data collection.

Although volume of down wood tends to be a measurement of down wood amount that also closely relates to wildlife use it is more complex to measure. Volume requires determining the length, the diameter of both ends of a log or use of a taper equation. Formulas are then used to calculate the volume using these variables. Taper equations will vary by species and decay class of down wood.

Number of down wood pieces per acre may offer the least amount of information when it comes to describing the amount of down wood in an area. Without knowing the length or diameters of the pieces, it has little value in describing wildlife habitat. As an example, 20 pieces of 10 foot logs 4 inches in diameter will have a much lower volume and percent cover than 20 pieces of 80 foot logs greater than 20 inches in diameter, thus resulting in different habitat conditions.

Linear feet of down wood is an often used measurement that also does not fully describe the amount of down wood in an area without knowing the diameters of the down wood being left. As an example, 500 linear feet of 4 inch diameter logs will have a much lower volume and percent cover than 500 linear feet of 20 inch diameter logs, thus resulting in different habitat conditions.

Whereas tons of logs per acre is a common measure used to describe fuel loadings, it is not as closely related to wildlife use and several assumptions and variables must be factored in to determining tons per acre. Weight of logs is based on estimated dry weight and will vary with species and decay class. An “average” species and decay class composition is usually used in tons per acre calculations based on vegetation type.

In order to compare data among studies, down wood data were converted into a common measure of percent cover. The forest inventory down wood data used in DecAID were used to calculate amount of down wood on each plot in terms of percent cover, volume, logs/ha, and linear meters. Regressions were then run on the data to calculate the relationships among 4 measures (see Table 1a-c). Correlations between percent cover and other measures were strongest with a small minimum diameter. Correlations between percent cover and logs/ha tended to be the weakest relationships. Based on the analysis, when data were reported in the literature using one of these other measures, we converted it to percent cover using the regression equations if the r2 value was > 0.75.

See the “Links to Useful Information” section of the Guide to the Interpretation and Use of the DecAID Advisor for tools to help with conversions to percent cover.

Table 1a. Summary of linear regressions for log volume (m3/ha), logs/ha, and linear meters/ha on percent cover of down wood > 12.5 cm (5 inches) diameter. The X variable is used in the formula to convert the various measures to percent cover (e.g., % cover = volume or linear meters/X).
Wildlife Habitat Type S-Class n r2 for Volume (m3/ha) X variable for Volume (m3/ha) r2 for Logs/ha X variable for Logs/ha r2 for Linear meters/ha X variable for Linear meters/ha
Eastside Mixed Conifer Forest Early 733 0.9158 23.54 0.8155 110.09 0.9536 467.57
Eastside Mixed Conifer Forest Mid 1089 0.9103 24.36 0.8177 103.96 0.9508 458.35
Eastside Mixed Conifer Forest Late 515 0.7917 29.78 0.8120 85.19 0.9385 398.78
Eastside Mixed Conifer Forest Mid & Late 1604 0.8373 26.58 0.8085 96.27 0.9422 433.94
Ponderosa Pine/Douglas-fir Early 353 0.4530 27.36 0.8023 132.40 0.9403 474.75
Ponderosa Pine/Douglas-fir Mid 757 0.8369 27.11 0.7825 128.09 0.9359 447.02
Ponderosa Pine/Douglas-fir Late 344 0.8443 29.67 0.7111 112.28 0.9289 411.51
Ponderosa Pine/Douglas-fir Mid & Late 1101 0.8383 28.03 0.7565 122.37 0.9321 434.17
Lodgepole Pine Early 431 0.9133 17.68 0.8082 106.71 0.9760 594.82
Lodgepole Pine Mid 174 0.7644 21.65 0.8038 97.24 0.9635 535.06
Montane Mixed Conifer Forest Early 473 0.8836 27.31 0.8147 107.90 0.9165 418.10
Montane Mixed Conifer Forest Mid 860 0.8812 26.37 0.8444 94.49 0.9348 435.94
Montane Mixed Conifer Forest Late 425 0.8976 31.47 0.8142 79.93 0.9344 363.30
Montane Mixed Conifer Forest Mid & Late 1285 0.8821 28.32 0.8297 88.92 0.9277 408.16
Southwest Oregon Mixed Conifer-Hardwood Forest Early 152 0.8463 36.47 0.7593 99.63 0.8576 354.72
Southwest Oregon Mixed Conifer-Hardwood Forest Mid 217 0.8684 34.02 0.7879 99.28 0.9076 366.92
Southwest Oregon Mixed Conifer-Hardwood Forest Late 137 0.9067 34.28 0.8312 86.82 0.9083 353.92
Southwest Oregon Mixed Conifer-Hardwood Forest Mid & Late 345 0.8838 34.13 0.7990 94.17 0.9076 361.6
Westside Lowland Conifer-Hardwood Forest Early 572 0.8777 31.38 0.8004 127.55 0.9186 376.13
Westside Lowland Conifer-Hardwood Forest Mid 817 0.8749 33.37 0.8094 104.37 0.9159 360.11
Westside Lowland Conifer-Hardwood Forest Late 656 0.9036 35.75 0.8134 79.85 0.9291 328.54
Westside Lowland Conifer-Hardwood Forest Mid & Late 1473 0.8897 34.62 0.7967 91.49 0.9202 343.52

Table 1b. Summary of linear regressions for log volume (m3/ha), logs/ha, and linear meters/ha on percent cover of down wood > 25 cm (10 inches) diameter. The X variable is used in the formula to convert the various measures to percent cover (e.g., % cover = volume or linear meters/X).

Wildlife Habitat Type S-Class n r2 for Volume (m3/ha) X variable for Volume (m3/ha) r2 for Logs/ha X variable for Logs/ha r2 for Linear meters/ha X variable for Linear meters/ha
Eastside Mixed Conifer Forest Early 643 0.9374 30.47 0.7993 55.56 0.9605 333.31
Eastside Mixed Conifer Forest Mid 979 0.9309 31.54 0.7994 53.06 0.9478 324.06
Eastside Mixed Conifer Forest Late 494 0.8122 37.08 0.7900 45.94 0.9480 293.91
Eastside Mixed Conifer Forest Mid & Late 1473 0.8561 34.13 0.7916 49.72 0.9457 309.93
Ponderosa Pine/Douglas-fir Early 448 0.5765 41.01 0.6982 59.56 0.9285 322.10
Ponderosa Pine/Douglas-fir Mid 925 0.7556 37.02 0.7042 56.86 0.9284 305.96
Ponderosa Pine/Douglas-fir Late 470 0.7737 37.85 0.7453 54.05 0.9435 296.21
Ponderosa Pine/Douglas-fir Mid & Late 1395 0.7630 37.36 0.7190 55.71 0.9341 301.96
Lodgepole Pine Early 659 0.8038 25.98 0.7830 45.92 0.9693 417.44
Lodgepole Pine Mid 273 0.8095 29.87 0.8317 50.65 0.9461 372.81
Montane Mixed Conifer Forest Early 951 0.9136 36.27 0.7816 52.87 0.9261 283.57
Montane Mixed Conifer Forest Mid 1504 0.9006 36.05 0.8562 51.33 0.9310 292.30
Montane Mixed Conifer Forest Late 903 0.9165 41.17 0.8377 44.66 0.9358 254.87
Montane Mixed Conifer Forest Mid & Late 2407 0.9059 38.70 0.8437 47.88 0.9288 272.93
Southwest Oregon Mixed Conifer-Hardwood Forest Early 213 0.8889 40.86 0.7445 54.57 0.9035 262.53
Southwest Oregon Mixed Conifer-Hardwood Forest Mid 311 0.8787 45.84 0.8006 51.04 0.9316 233.97
Southwest Oregon Mixed Conifer-Hardwood Forest Late 193 0.9374 44.16 0.7977 42.61 0.9403 237.74
Southwest Oregon Mixed Conifer-Hardwood Forest Mid & Late 504 0.9009 45.14 0.7936 47.52 0.9352 235.54
Westside Lowland Conifer-Hardwood Forest Early 1168 0.9251 44.48 0.7674 62.04 0.9441 231.94
Westside Lowland Conifer-Hardwood Forest Mid 1644 0.9120 43.31 0.8163 57.63 0.9407 238.92
Westside Lowland Conifer-Hardwood Forest Late 1442 0.9455 44.96 0.8473 45.74 0.9573 230.59
Westside Lowland Conifer-Hardwood Forest Mid & Late 3090 0.9317 44.28 0.8197 50.63 0.9498 234.02

Table 1c. Summary of linear regressions for log volume (m3/ha), logs/ha, and linear meters/ha on percent cover of down wood > 50 cm (20 inches) diameter. The X variable is used in the formula to convert the various measures to percent cover (e.g., % cover = volume or linear meters/X).

Wildlife Habitat Type S-Class n r2 for Volume (m3/ha) X variable for Volume (m3/ha) r2 for Logs/ha X variable for Logs/ha r2 for Linear meters/ha X variable for Linear meters/ha
Eastside Mixed Conifer Forest Early 296 0.9629 49.91 0.5143 24.90 0.9687 193.61
Eastside Mixed Conifer Forest Mid 476 0.9564 49.44 0.6273 25.74 0.9584 186.05
Eastside Mixed Conifer Forest Late 313 0.8116 58.20 0.7031 19.94 0.9437 177.17
Eastside Mixed Conifer Forest Mid & Late 789 0.8512 54.48 0.6485 22.40 0.9496 180.94
Ponderosa Pine/Douglas-fir Early 217 0.5487 74.80 0.5544 28.39 0.9309 172.99
Ponderosa Pine/Douglas-fir Mid 422 0.7306 57.87 0.6060 25.37 0.9404 178.67
Ponderosa Pine/Douglas-fir Late 258 0.7121 56.56 0.6680 24.40 0.9484 181.46
Ponderosa Pine/Douglas-fir Mid & Late 680 0.7229 57.32 0.6288 24.97 0.9437 179.82
Lodgepole Pine Early 184 0.5812 60.19 0.5603 27.38 0.9231 191.90
Lodgepole Pine Mid 106 0.8135 63.09 0.7798 25.11 0.9271 163.27
Montane Mixed Conifer Forest Early 602 0.9512 54.93 0.7398 24.27 0.9710 167.49
Montane Mixed Conifer Forest Mid 932 0.9150 55.03 0.7731 26.39 0.9637 167.14
Montane Mixed Conifer Forest Late 700 0.9528 57.58 0.8107 20.56 0.9637 162.85
Montane Mixed Conifer Forest Mid & Late 1632 0.9384 56.61 0.7793 22.78 0.9636 164.48
Southwest Oregon Mixed Conifer-Hardwood Forest Early 108 0.9508 58.41 0.6535 27.84 0.9572 157.36
Southwest Oregon Mixed Conifer-Hardwood Forest Mid 183 0.8852 60.75 0.6963 25.12 0.9707 153.33
Southwest Oregon Mixed Conifer-Hardwood Forest Late 126 0.9566 57.43 0.7427 17.21 0.9660 161.69
Southwest Oregon Mixed Conifer-Hardwood Forest Mid & Late 309 0.9110 59.37 0.6855 21.83 0.9680 156.81
Westside Lowland Conifer-Hardwood Forest Early 783 0.9516 59.71 0.7376 28.39 0.9651 152.48
Westside Lowland Conifer-Hardwood Forest Mid 1170 0.9418 58.96 0.7836 27.97 0.9687 154.60
Westside Lowland Conifer-Hardwood Forest Late 1164 0.9623 58.61 0.7693 21.45 0.9741 157.64
Westside Lowland Conifer-Hardwood Forest Mid & Late 2334 0.9540 58.75 0.7633 24.03 0.9719 156.44

Several examples of down wood of different sizes and what they represent in terms of percent cover are given in Table 2. The conversions from log size to percent cover use the formula: % cover = (((((LED+SED)/2)/12)*L)/43560)*100; where LED=large end diameter (in), SED=small end diameter (in), and L=length (ft). Small end diameter was calculated based on the "rule of thumb" of 1 inch taper per 8 feet of length (Ken Cormier, pers. comm.), and the formula: SED = LED-(L/8).

Table 2. Conversion of individual pieces of down wood to percent cover given large end diameter and length, assuming 1 inch of taper per 8 feet of length.

Large end diameter (in) Down wood length (ft) % cover on per acre basis
6 20 0.018
8 30 0.035
10 40 0.057
12 20 0.041
12 40 0.073
12 65 0.099
15 20 0.053
15 40 0.096
15 60 0.129
15 75 0.148
20 20 0.072
20 40 0.134
20 60 0.187
20 80 0.230
20 100 0.263
30 20 0.110
30 40 0.210
30 60 0.301
30 80 0.383
30 100 0.454
30 140 0.569
40 20 0.148
40 40 0.287
40 60 0.416
40 100 0.646
40 160 0.918

If trees are to be felled to provide down wood, or left standing to provide down wood at a later time, percent cover contribution of each tree can be estimated using taper equations. Clicking on the wildlife habitat type and subregion below will return a table with rough estimates of percent of an acre covered by different species of trees of given dbh when converted to down wood. The Flewelling Profile Model (Volume Estimator Library, Forest Vegetation Simulator) was used to determine height of a tree to a 5 inch top (12.7 cm). Total height of trees is used in the equations; this variable was estimated by averaging tree height by species and dbh class from inventory data specific to each Wildlife Habitat Type and subregion. These data were not updated with the latest inventory data for version 3.0 because the relationship between tree dbh and height should not have changed between the 2 old data and new data. The midpoint dbh for each class was input to the model as the dbh. A different version of the Flewelling Profile Model was used for each tree species.

Westside Lowland Conifer-Hardwood Forest - WA Coast

Westside Lowland Conifer-Hardwood Forest - OR Coast

Westside Lowland Conifer-Hardwood Forest - WA W Cascades

Westside Lowland Conifer-Hardwood Forest - OR W Cascades

Southwest Oregon Mixed Conifer-Hardwood Forest

Montane Mixed Conifer Forest

Eastside Mixed Conifer Forest - N Cascades/Rocky Mtns.

Eastside Mixed Conifer Forest - E Cascades/Blue Mtns.

Ponderosa Pine/Douglas-fir Forest

Table 3. Percent of an acre covered by different species of tree of given dbh when the tree is felled to provide down wood for the Westside Lowland Conifer-Hardwood Forest - WA Coast. Blank cells indicate insufficient data were available on tree heights for that species and dbh class.

DBH (in) Douglas-fir % cover Western redcedar % cover Western hemlock % cover
10 0.066 0.054 0.062
15 0.123 0.098 0.127
20 0.204 0.161 0.212
25 0.295 0.255 0.312
30 0.400 0.319 0.421
35 0.536 0.431 0.535
40 0.628 0.508 0.639
50 0.871 0.692 0.880
60 1.169 0.921 1.050

Table 4. Percent of an acre covered by different species of tree of given dbh when the tree is felled to provide down wood for the Westside Lowland Conifer-Hardwood Forest - OR Coast. Blank cells indicate insufficient data were available on tree heights for that species and dbh class.

DBH (in) Douglas-fir % cover Western redcedar % cover Western hemlock % cover
10 0.071 0.052 0.065
15 0.138 0.100 0.128
20 0.239 0.163 0.210
25 0.359 0.227 0.312
30 0.489 0.301 0.422
35 0.594 0.389 0.541
40 0.715 0.418 0.662
50 0.908 0.672 0.878
60 1.102 0.878
70 1.401 1.038
80 1.583

Table 5. Percent of an acre covered by different species of tree of given dbh when the tree is felled to provide down wood for the Westside Lowland Conifer-Hardwood Forest - WA W Cascades. Blank cells indicate insufficient data were available on tree heights for that species and dbh class.

DBH (in) Douglas-fir % cover Western redcedar % cover Western hemlock % cover
10 0.074 0.054 0.062
15 0.139 0.106 0.127
20 0.229 0.177 0.214
25 0.332 0.259 0.300
30 0.439 0.340 0.396
35 0.535 0.426 0.503
40 0.685 0.538 0.590
50 0.824 0.715 0.770
60 1.032 0.936
70 1.290 0.999
80 1.464 1.254

Table 6. Percent of an acre covered by different species of tree of given dbh when the tree is felled to provide down wood for the Westside Lowland Conifer-Hardwood Forest - OR W Cascades. Blank cells indicate insufficient data were available on tree heights for that species and dbh class.

DBH (in) Douglas-fir % cover Western redcedar % cover Western hemlock % cover
10 0.069 0.055 0.059
15 0.137 0.104 0.125
20 0.232 0.173 0.213
25 0.335 0.261 0.310
30 0.445 0.350 0.409
35 0.546 0.415 0.518
40 0.657 0.494 0.590
50 0.827 0.707 0.831
60 1.045 0.894 1.046
70 1.274
80 1.447

Table 7. Percent of an acre covered by different species of tree of given dbh when the tree is felled to provide down wood for the Southwest Oregon Mixed Conifer-Hardwood Forest. Blank cells indicate insufficient data were available on tree heights for that species and dbh class.

DBH (in) Douglas-fir % cover Ponderosa pine % cover Western hemlock % cover White fir % cover
10 0.061 0.045 0.054 0.061
15 0.120 0.104 0.117 0.125
20 0.203 0.191 0.191 0.212
25 0.298 0.284 0.291 0.306
30 0.398 0.403 0.396 0.405
35 0.489 0.522 0.460 0.498
40 0.592 0.767 0.577
50 0.740 0.839 0.810
60 0.951 1.094
70 1.139
80 1.394

Table 8. Percent of an acre covered by different species of tree of given dbh when the tree is felled to provide down wood for the Montane Mixed Conifer Forest. Blank cells indicate insufficient data were available on tree heights for that species and dbh class.

DBH (in) Douglas-fir % cover Western hemlock % cover Mountain hemlock % cover Western redcedar % cover Engelmann spruce % cover Western larch % cover White fir/Grand fir % cover Subalpine fir % cover Western white pine % cover Lodgepole pine % cover
10 0.056 0.053 0.045 0.048 0.064 0.076 0.056 0.055 0.055 0.064
15 0.109 0.113 0.096 0.098 0.122 0.139 0.113 0.104 0.113 0.113
20 0.180 0.193 0.160 0.154 0.194 0.212 0.187 0.159 0.179 0.164
25 0.261 0.287 0.231 0.234 0.271 0.283 0.266 0.214 0.269 0.202
30 0.347 0.379 0.304 0.316 0.352 0.353 0.346 0.259 0.333 0.291
35 0.425 0.471 0.372 0.383 0.441 0.444 0.461 0.319 0.413
40 0.536 0.576 0.464 0.491 0.512 0.489 0.500 0.330 0.543
50 0.667 0.747 0.617 0.677 0.661 0.630 0.760
60 0.831 0.886 0.822 0.778
70 1.146 0.953
80 1.291

Table 9. Percent of an acre covered by different species of tree of given dbh when the tree is felled to provide down wood for the Eastside Mixed Conifer Forest - N Cascades/Rocky Mtns. Blank cells indicate insufficient data were available on tree heights for that species and dbh class.

DBH (in) Douglas-fir % cover Western hemlock % cover Mountain hemlock % cover Western redcedar % cover Engelmann spruce % cover Western larch % cover White fir/Grand fir % cover Subalpine fir % cover Western white pine % cover Lodgepole pine % cover
10 0.058 0.051 0.082 0.071 0.062 0.070 0.057 0.077 0.069 0.080
15 0.108 0.105 0.146 0.133 0.117 0.131 0.102 0.140 0.126 0.131
20 0.172 0.171 0.222 0.212 0.183 0.207 0.159 0.217 0.185 0.208
25 0.240 0.252 0.292 0.297 0.267 0.304 0.231 0.301 0.313
30 0.309 0.330 0.376 0.361 0.303 0.418 0.306 0.344
35 0.351 0.410 0.444 0.467 0.361
40 0.407 0.499 0.564 0.545 0.487
50 0.486 0.738
60 0.798

Table 10. Percent of an acre covered by different species of tree of given dbh when the tree is felled to provide down wood for the Eastside MIxed Conifer Forest - E Cascades/Blue Mtns. Blank cells indicate insufficient data were available on tree heights for that species and dbh class.

DBH (in) Douglas-fir % cover Ponderosa pine % cover Western larch % cover Engelmann spruce % cover White fir/Grand fir % cover Lodgepole pine % cover
10 0.052 0.046 0.076 0.066 0.051 0.066
15 0.101 0.099 0.140 0.126 0.102 0.117
20 0.166 0.171 0.223 0.203 0.171 0.173
25 0.241 0.253 0.304 0.289 0.246 0.217
30 0.316 0.335 0.388 0.357 0.329
35 0.380 0.420 0.477 0.474 0.411
40 0.454 0.506 0.582 0.549 0.477
50 0.582 0.678 0.690 0.809 0.628
60 0.731 0.786

Table 11. Percent of an acre covered by different species of tree of given dbh when the tree is felled to provide down wood for the Ponderosa Pine/Douglas-fir Forest. Blank cells indicate insufficient data were available on tree heights for that species and dbh class.

DBH (in) Douglas-fir % cover Ponderosa pine % cover Lodgepole pine % cover
10 0.045 0.041 0.056
15 0.085 0.086 0.104
20 0.139 0.152 0.164
25 0.208 0.228 0.233
30 0.262 0.308 0.335
35 0.335 0.385
40 0.449 0.469
50 0.630 0.660
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